Education:

1998-2004: Doctor of Veterinary Medicine, Hokkaido University School of Veterinary Medicine, Japan 

2004-2008: Doctorate, Hokkaido University Graduate School of Medicine, Japan (Advisor: Toshiyuki Sawaguchi and Kikuro Fukushima)

Professional Experience:

2020.3-present: Principal Investigator, Chinese Institute for Brain Research, Beijing, China 

2019.3-present: Visiting Scientist, Department of Otorhinolaryngology: Head and Neck Surgery, University of Pennsylvania School of Medicine, USA

2019.3-2020.2: Research Assistant Professor, Department of Veterinary Medicine, Iwate University, Japan

2015.7-2019.2: Research Associate, Department of Otorhinolaryngology: Head and Neck Surgery, University of Pennsylvania School of Medicine, USA (Advisor: Steven J. Eliades)

2009.4-2015.6: Research Associate, Department of Otorhinolaryngology: Head and Neck Surgery, University of Pennsylvania School of Medicine, USA (Advisor: Yale E. Cohen)

2008.9-2009.3: Postdoctoral Research Fellow, Department of Psychological and Brain Sciences, Center for Cognitive Neuroscience, Dartmouth College, USA (Advisor: Yale E. Cohen)

2008.4-2008.8: Postdoctoral Research Fellow, Department of Psychology, Tokyo Metropolitan Institute for Neuroscience, Japan (Advisor: Masataka Watanabe)